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It functions as the Alt key in Unix and Windows environments. It serves the function of the solid-Apple key in Apple II applications.

Like the Shift and Control keys, the Option key serves as a modifier for the Command key shortcuts, as well as being used to type many special characters. The Option key ( ⌥), for entering diacritics and other special characters.The open-Apple key was combined with the Command key on Apple Desktop Bus keyboards (which were used on both the Apple IIgs and several years of Macintosh models) where it remained after the Apple II line was discontinued. The "open" (hollow) and separate "closed" (solid) Apple logo keys on the Apple II series, served functions similar to that of the Command key.Compared to their equivalents on the standard IBM PC keyboard layout the Command key and the Option key are located in reverse order. The key functions as a Meta key or Super key in Unix-like environments, and is equally equivalent to the Windows key in Windows environments, although in common applications it performs the same function as the Windows Control key. The Command key ( ⌘), used in most Mac keyboard shortcuts.Features different from other keyboards include: Some of these keys have unique symbols defined in the Unicode block Miscellaneous Technical.

To serve the functionality of the Macintosh operating systems (and because of historical differences), the Apple Keyboard's layout differs somewhat from that of the ubiquitous IBM PC keyboard, mainly in its modifier and special keys.